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Checklist of Onthophagus of Russia and adjacent lands
©Composed by Andre Gorodinski, 2006.
  SPECIES DISTRIBUTION SYNONYM
  Onthophagus acrisius Balthasar, 1946 China (Fujian)  
  Onthophagus acuticollis Gillet, 1927 Taiwan  
  Onthophagus afghanus Petrovitz , 1961 Kirgizstan,Turkmenistan,Uzbekistan, Tadzhikistan,
Afganistan		  
  Onthophagus agilis Matsumura, 1938 Taiwan  
  Onthophagus akinini Koenig, 1889 Turkmenistan, Uzbekistan  
  Onthophagus amyntas Olivier, 1789 Russia, Azerbaidzhan, Armenia, Georgia, Middle
Asia,China, Afganistan, Mediterranean 		Onthophagus tages, Olivier 1789; Onthophagus
huebneri, Fabricius, 1792
  Onthophagus amplexus Sharp, 1875 China,India  
  Onthophagus andalusicus andalusicus Waltl,
1835 		Russia ,Ukraina, Kazahstan, S.E. Europe, N. Afrika,
Iran		 Onthophagus maurus Lucas, Onthophagus
tescuorum Semenov & Medvedev ,1927
  Onthophagus andalusicus persianus Olsoufiev,
1900		 Georgia, Azerbaidzhan, Armenia, Iran  
  Onthophagus angorensis Petrovitz , 1963 Azerbaidzhan, Armenia, Georgia,Turkey,
Turkmenistan, Iran		  
  Onthophagus anguicorius Boucomont, 1914 China (Hongkong, Hainan),Vietnam, Taiwan, Laos  
  Onthophagus anguliceps Boucomont, 1914 China (Xizang), India, Taiwan, Bhutan, Burma  
  Onthophagus argyropygus Gillet,1927 China (mainland of China , Hongkong), Japan,
Taiwan		  
  Onthophagus armatus Blanchard, 1853 China (Fujian, Guangdong), Vietnam, Taiwan,
Cambodia, Burma, Malaysia		  
  Onthophagus arnoldii Kabakov , 1982 Turkmenistan,Tadzhikistan  
  Onthophagus ater Waterhouse, 1875 China (Fujian,Guizhou, Sichuan), Japan,
Taiwan,Korea		 Onthophagus granulipennis Frivaldsky, 1892;
Onthophagus chuzenjianus, -kawarinus
Matsumura et Yohena, 1939
  Onthophagus atramentarius Ménétriès, 1832 Russia (Caucasus) ,Georgia, Azerbaidzhan, Turkey,
Greece, Iran, Irak, Egypt 		Onthophagus orcas Ménétriès, 1832 ;
Onthophagus felscheri Reitter , 1891;
Onthophagus peropacus Kirschenblatt ,1935
  Onthophagus atripennis Waterhouse, 1875 Russia (Far East), China(Sichuan, Beijing, Fujian),
Japan, Korea		 Onthophagus cicatricosus Balthasar, 1935;
Onthophagus akirai, -ibonus, -kogatamus,
-shigeoi Matsumura et Yohema, 1937
  Onthophagus balthasari Vsetecka,1939 China, SE. China  
  Onthophagus basicruentatus Goidanich, 1926 NW. China (Fujian), Japan Onthophagus oshimanus Nakane,1960
  Onthophagus basipustulatus Heyden, 1889 Kazahstan, Kirgizstan,Uzbekistan, Afganistan  
  Onthophagus bivertex Heyden,1887 Russia (Siberia), China (Beijing,Shanxi, Fujian,
Sichuan), Japan,Korea		 Onthophagus ibex Motschulsky, 1854; Onthophagus mandarinus Harold, 1859;
Onthophagus minokuchianus Matsumura, 1934;
Onthophagus kozunonis, -oniellus, -yohenai
Matsumura, 1937
  Onthophagus boucomontianus Balthasar, 1935 China (Yunnan), Vietnam  
  Onthophagus brutus Arrow, 1931 China (Yunnan), Vietnam, Laos, Thailand  
  Onthophagus carinensis Boucomont, 1914 China (Yunnan), Burma  
  Onthophagus carpanetoi Pittino, 1982 Turkey,Irak ,Syria  
  Onthophagus cernyi Balthasar, 1935 China (Yunnan)  
  Onthophagus chinensis Balthasar, 1952 China (Fujian)  
  Onthophagus chokakurinus Matsumura, 1938 Taiwan  
  Onthophagus clitellifer Reitter, 1894 Russia (Siberia), China (Gansu, Liaonirig, Beijing),
Mongolia		 Onthophagus transbaicalicus Csiki, 1901
  Onthophagus coenobita Herbst , 1783 W.Russia ,Ukraina, Europe, Georgia, Azerbaidzhan,
Armenia, Syria,W. Iran		 Onthophagus fulgens Brahm , 1790 ;
Onthophagus tenuicornis Preyssler, 1790 ;
Onthophagus rufipes Ménétriès,1832 ;
Onthophagus fulvipes Faldermann, 1835 ;
Onthophagus subprominulus ,-tricuspis Mulsant,
1842
  Onthophagus collaris Kabakov China (Yunnan)  
  Onthophagus conspersus Reitter, 1893 Azerbaidzhan,Turkmenistan, Uzbekistan, Tadzhikistan,
Afganistan, N. Iran		  
  Onthophagus convexicollis Boheman, 1858 China (Hongkong)  
  Onthophagus coracinoides Kabakov, 1994 China (Yunnan), Vietnam  
  Onthophagus coracinus Boucomont, 1914 China (Fujian, Hongkong, Yunnan), Vietnam, Burma,
Laos		  
  Onthophagus crassicollis Boucomont, 1913 China (Yunnan), Vietnam  
  Onthophagus cruciatus Menetries, 1832 Azerbaidzhan, Armenia, Georgia,Turkey,Irak ,W.Iran Onthophagus centromaculatus Redtenbacher,
1843
  Onthophagus curvispina Reitter, 1893 China (Beijing)  
  Onthophagus dama Fabricius, 1798 China, India  
  Onthophagus dapcauensis Boucomont, 1921 China (Guangdong, Hongkong), Vietnam, Taiwan  
  Onthophagus deflexicolis Lansberge, 1883 China (Guizhou, Sichuan), Thailand, Laos, Vietnam,
Malaysia		  
  Onthophagus diabolicus Harold, 1877 China (Yunnan) India, Laos, Vietnam, Malaysia,
Indonesia 		 
  Onthophagus discedens Sharp, 1875 China (Sichuan, Yunnan), Burma, Laos, Sikkim,
Bangladesh, Indonesia, Japan		  
  Onthophagus dissentaneus Balthasar, 1964 China (Yunnan)  
  Onthophagus diversicornis Kirschenblatt, 1935 Azerbaidzhan, Armenia  
  Onthophagus dorbignyi Olsouffiev, 1900 S.Turkmenistan, Iran, Afganistan  
  Onthophagus dorsofasciatus Fairmaire, 1893 China (Shandong), Vietnam  
  Onthophagus dorsosignatus d’Orbigny , 1898 Azerbaidzhan, Armenia, Turkey, N. Iran , Irak ,Syria  
  Onthophagus dubernardi Boucomont, 1914 China (Hubei, Guizhou, Sichuan, Yunnan, Xizang),
Vietnam, Burma, Sikkim		  
  Onthophagus duporti Boucomont, 1914 China (Yunnan), Vietnam  
  Onthophagus egenus Harold, 1877 China (Hongkong), Indonesia  
  Onthophagus egurianus Matsumura, 1938 Taiwan  
  Onthophagus falzonii Goidanich, 1926 Turkey,Irak ,Syria ,Iran  
  Onthophagus finschi Harold, 1877 Kazahstan, Kirgizstan, N. China, Mongolia carpeolus Ballion, 1878 ; protzenkoi,
paraprotzenkoi Tesar, 1967
  Onthophagus fissicornis Steven , 1809 Russia (Caucasus),S. Ukraina, Georgia, Azerbaidzhan,
Armenia, Turkey, S.E. Europe, S.Turkmenistan, Iran,
Irak, Syria		  
  Onthophagus flagrans Reitter, 1893 Turkmenistan, Uzbekistan, Tadzhikistan, Kazahstan,
Kirgizstan, N. Iran , Armenia, Afganistan		  
  Onthophagus fodiens Waterhouse, 1875 Russia (Far East), China (Hubei, Shanxi, Jiangsu,
Jiangxi, Zhejiang, Fujian, Sichuan), Japan,Korea		 Onthophagus hikosanus, -sabosanus, -ushiodai,
yubarinus Matsumura et Yohena, 1937
  Onthophagus formaneki Reitter, 1897 Azerbaidzhan, Armenia, Turkey, Syria Onthophagus bifenestratus Reitter, 1898
  Onthophagus formosanus Gillet, 1924 Taiwan  
  Onthophagus fortigibber Reitter, 1909 E. Georgia, W. Azerbaidzhan, E. Turkey Onthophagus tricuspis Semenov , 1899 ; Onthophagus unxovi Bogachev , 1930
  Onthophagus fracticornis Preyssler, 1790 Russia, Europe, Caucasus, Turkey, Kazahstan,
Turkmenistan, Iran, N. Afrika, Syria		 Onthophagus herbsti Brahm., 1790 ; Onthophagus
assimilis Hoppe , 1795; Onthophagus irroratus
Falderman , 1835; Onthophagus panperatus, -
sublaminatus,- subrecticornis,- tricuspidatus
Mulsant, 1842; Onthophagus anonimus Delabie,
1956
  Onthophagus funebris Boucomont, 1920 China (Yunnan), Vietnam  
  Onthophagus furcatus Fabricius , 1781 Russia, Caucasus, Europe, Ukraina, N.Kazahstan,
Turkmenistan, Uzbekistan, Iran		 Onthophagus vitulus Laicharting , 1781;
Onthophagus terminatum Falderman , 1835 ;
Onthophagus fossatus Castelnau , 1840 ;
Onthophagus bicornutus ,- bidentarus,- laminiger
Mulsant,1842; Onthophagus mnizhechi
Hochenwarth,1873
  Onthophagus furciceps Marseul , 1869 Moldavia, Ukraina, S.E. Europe, Turkey, Iran, Irak,
Syria, Lebanon		 Onthophagus kindermanni Harold ,1877 ;
Onthophagus strabo Reitter ,1893
  Onthophagus furcicornis Reitter , 1893 S. Russia, Moldavia, Ukraina, Georgia, Azerbaidzhan,
Armenia, Turkey, Syria, Lebanon		 Onthophagus parmatus Reitter , 1893 ;
Onthophagus citellorum Medvedev, 1965
  Onthophagus fuscopunctatus Boucomont, 1914 China (Guangdong, Hongkong), Vietnam, Burma,
Laos, Malaysia, Celebes, E.Asia		  
  Onthophagus gagates Hope, 1831 China (Sichuan, Yunnan, Xizang), India, Kashmir,
Vietnam, Russia (Siberia)		 Onthophagus angulatus Redtenbacher, 1848
  Onthophagus gibbosus gibbosus Scriba, 1790 Russia, Caucasus, Mongolia, China , Asia Minor,
Turkey, Europe		 Onthophagus juvensis Scriba, 1790; Onthophagus
tages,-dubius, -difformis, -unituberculatus,
-sycophanta, -umbrinus Mulsant, 1842;
Onthophagus huebneri Erichson, 1847;
Onthophagus amynthus Mulsant et Rey, 1871;
Onthophagus ganglbaueri Reitter, 1891
  Onthophagus gibbosus koshantschikoffi Reitter,
1891		 Middle Asia, Iran, Afganistan, W. China  
  Onthophagus gibbosus schnabeli Splichal, 1910 Afganistan, N. Pakistan, N.W. India  
  Onthophagus gibbulus gibbulus Pallas, 1781 Russia, Europe, Middle Asia, China, Mongolia, Korea,
Japan		 Onthophagus austriacus Panzer, 1793
  Onthophagus gibbulus laticornis Gebler, 1823 Russia (S.E. Siberia), Mongolia, N. China  
  Onthophagus gibbulus rostrifer Reitter, 1893 Transcaucasus, E. Turkey, N. Iran  
  Onthophagus ginyunensis Vsetecka, 1942 China (Zhejiang)  
  Onthophagus glasunovi Koshantschikov, 1894 Uzbekistan, Tadzhikistan  
  Onthophagus gorodinskii Kabakov sp.n. in litt China (Sichuan)  
  Onthophagus gracilipes Boucomont, 1914 China (Yunnan), Burma, Laos, Vietnam  
  Onthophagus grossepunctatus Reitter, 1905 Russia , Ukraina, Europe, Azerbaidzhan, Armenia,
Georgia, Turkey 		 
  Onthophagus hajimei Masumoto, 1984 Taiwan  
  Onthophagus haroldi Ballion, 1871 Turkmenistan, Kazahstan,Uzbekistan, Kirgizstan,
Tadzhikistan, N.E. Iran , N.Afganistan, N. China		  
  Onthophagus hastifer Lansberge, 1885 China (Guangdong), Vietnam, Thailand, Burma,
Taiwan		  
  Onthophagus heyrovskyi Vsetecka, 1944 China (Sichuan)  
  Onthophagus hirsutulus Lansberge, 1883 China, Taiwan, Indonesia, Philippines  
  Onthophagus hissariensis Kabakov , 2006 Tadzhikistan  
  Onthophagus horni Balthasar, 1935 China  
  Onthophagus(Indachorius) hsui Masumoto et
Chen et Ochi, 2004		 Taiwan  
  Onthophagus hvangheus Stebnicka , 1973 N. Korea  
  Onthophagus illyricus Scopoli, 1763 Russia, Turkmenistan, Europe, Turkey, Iran, Syria Onthophagus urus Ménétriès, 1832 ; Onthophagus
menetriesi Faldermann , 1835; Onthophagus
brisouti d’Orbigny , 1897
  Onthophagus inae Balthasar, 1963 Taiwan  
  Onthophagus inelegans Balthasar, 1935 China (Yunnan, Sichuan)  
  Onthophagus japonicus Harold, 1874 Russia (Far East), China (Shaanxi, Guizhou), Japan,
Korea		  
  Onthophagus joannae Goljan, 1953 W.Ukraina, Europe  
  Onthophagus kentingensis Nomura, Taiwan  
  Onthophagus klapperichi Balthasar, 1954 China, Taiwan  
  Onthophagus kleinei Balthasar, 1935 China  
  Onthophagus komareki Balthasar, 1935 China  
  Onthophagus konoi Matsumura, 1938 Taiwan  
  Onthophagus kozlovi Kabakov,1990 China (Xiahe, Gansu, Sichuan), E.Tibet  
  Onthophagus kuatunensis Balthasar, 1942 China (Yunnan)  
  Onthophagus kukunorensis Kabakov, 1990 China(N.E. Tibet)  
  Onthophagus kulti Balthasar, 1952 China (Fujian)  
  Onthophagus kuluensis Bates, 1891 China (Xizang), India, Kashmir  
  Onthophagus laevis Harold, 1880 China (Hubei, Yunnan, Xizang), Burma, Laos, Malaysia,
Japan, Indonesia		  
  Onthophagus laevis ssp. asiaticus Boucomont,
1919		 China (Yunnan)  
  Onthophagus laevis ssp. lampromelas Faiimaire,
1891		 China  
  Onthophagus lamellatus Boucomont, 1924 China; India Onthophagiis laminatus Taschenb.1879
  Onthophagus laotianus Boucomont, 1914 China (Yunnan), N.Vietnam  
  Onthophagus laticornis Gebler ,1823 Russia (Siberia), Mongolia, N. China  
  Onthophagus lemur Fabricius , 1781 Russia, Europe, Ukraina, Georgia, Azerbaidzhan,
Armenia, Lebanon ,Syria, N.Iran		 Onthophagus quadrituberculatus Laicharting ,
1781; Onthophagus decempustulatus Schaller,
1783 ; Onthophagus lineolatus ,- egenus Mulsant,
1848 ; Onthophagus plurimacula Depoli, 1924 ;
Onthophagus simplicithorax Endrodi , 1955
  Onthophagus lenzi Harold, 1874 China (Lianong, Beijing, Henan, Shanxi, Jiangsu,
Jiangxi, Zhejiang, Fujian, Gjizhou, Sichuan), Japan,
Korea, N. India, Taiwan		 Onthophagus kuraruanus Matsumura, 1938
  Onthophagus leucomelas Solsky,1876 Turkmenistan, Uzbekistan, Tadzhikistan, Kazahstan,
Kirgizstan, N. Iran , Afganistan, N. China		  
  Onthophagus leucostigma Stevens, 1811 S.W. Russia , S. Ukraina, Kazahstan  
  Onthophagus limbatus Harold, 1789 China, Vietnam, Laos, Malaysia, Indonesia,
Calemantan		  
  Onthophagus lucidus Sturm , 1800 Russia, Ukraina, Europe, Georgia, Azerbaidzhan,
Armenia, Turkey, Europe, Irak, Syria, W. Iran		 Onthophagus nitidulus Brulle , 1832;
Onthophagus aequalis Endrodi ,1955
  Onthophagus lunatus Harold, 1868 China (Fujian, Guangdong, Hongkong), Vietnam,
Thailand, Cambodia		  
  Onthophagus luridipennis Boheman, 1858 China (Fujian, Hongkong, Yunnan ), India, E. Asia,
Burma, Vietnam		  
  Onthophagus lutosopictus Fairmaire, 1897 Japan  
  Onthophagus magnini Paulian, 1933 China (Yunnan), Vietnam  
  Onthophagus manipurensis Arrow, 1907 China (Xizang), India, Burma, Indonesia  
  Onthophagus marani Vsetecka, 1942 China (Zhejiang)  
  Onthophagus marginalis marginalis Gebler,
1817		 Russia (Siberia), W. Kazahstan, NE. China, Mongolia,
Korea		  
  Onthophagus marginalis marmoratus Ménétriès,
1832		 C. & E. Transcaucasus, Iran, C. & N. Afganistan,
S. Middle Asia		  
  Onthophagus marginalis nigrimargo Goidanich, 1926 - China (S. Tibet), N. India, S.
Afganistan, N. Pakistan		  
  Onthophagus marginalis przewalskii Kabakov,
2006		 W. China, E. Kazahstan, E. Kirgizstan  
  Onthophagus marmotae Kabakov , 1990 Uzbekistan, Tadzhikistan,Afganistan  
  Onthophagus masumotoi Kim, 1985 Taiwan  
  Onthophagus medvedevi Kabakov , 1982 Kazahstan,Kirgizstan, Tadzhikistan  
  Onthophagus mendicus Gillet,1924 Taiwan  
  Onthophagus missor Balthasar, 1938 Taiwan  
  Onthophagus miyabei Matsumura, 1937 China, NE. China  
  Onthophagus miyakei Ochi and Araya, 1992 Taiwan, Okinawa  
  Onthophagus mindolianus Matsumura, 1938 Taiwan  
  Onthophagus mulleri Lansberge, 1883 China (Yunnan), Vietnam  
  Onthophagus mushensis Matsumura, 1938 Taiwan  
  Onthophagus nagasawai Matsumura, 1938 Taiwan  
  Onthophagus nakatomii Matsumura, 1937 NE. China  
  Onthophagus naviculifer Boucomont, 1919 China (Yunnan), Laos  
  Onthophagus neofurcatus Godanich, 1926 China (Fujian,Jiangxi) Onthophagus furcatoides Reitter,1893
  Onthophagus nikolajevi Kabakov , 2006 China ( Sichuan , Gansu)  
  Onthophagus nitidiceps Fairmaire, 1893 China (Shanghai), Burma, Vietnam, Malaysia  
  Onthophagus nitidus nitidus Waterhouse, 1875 China (Jilin, Shanghai, Hongkong), Japan, Taiwan  
  Onthophagus novaki Boucomont , 1927 Syria,Israel,Lebanon muelleri Novak , 1921
  Onthophagus nuchicornis Linnaeus, 1758 Russia, China, Europe, C. & E. Asia Onthophagus planicornis Herbst, 1785;
Onthophagus acornis Geoffroy, 1785;
Onthophagus xiphias Fabricius, 1792;
Onthophagus trituberculatus Schrank, 1798;
Onthophagus alpinus Kolenati; Onthophagus
kolenatii Reitter, 1893; Onthophagus rubripes
Mulsant, 1842
  Onthophagus obliviosus Balthasar, 1935 China  
  Onthophagus ohbayashii Nomura, 1939 China, Japan,Korea  
  Onthophagus olsoufieffi Boucomont, 1924 Russia (Siberia), China (Gansu, Liaoning, Hubei,
Shandong), Japan,Korea, Mongolia		 Onthophagus penicillatus Olsouffiev, 1906;
Onthophagus kaudai Matsumura. 1937;
Onthophagus uedanus Matsumura, 1937
  Onthophagus opacicollis Reitter , 1893 Russia (Caucasus), Georgia, Azerbaidzhan, Armenia,
Europe , N. Afrika, Turkey, Iran, Syria		 Onthophagus schatzmayri Pierroti , 1959
  Onthophagus orientalis Harold, 1868 China (Fujian, Guangzhou, Hongkong, Guangxi),
India, Burma, Sikkim, Thailand, Cambodia, Laos,
Thailand, Malaysia, E. Asia		  
  Onthophagus osellai Pittino, 1982 Armenia, Turkey, W.Iran  
  Onthophagus ovatus Linnaeus , 1767 Russia, Ukraina, Europe, Kazahstan, Azerbaidzhan,
Armenia, Georgia, Turkey, Turkmenistan, Kazahstan,
Uzbekistan		 Onthophagus fucatus Mulsant, 1842;
Onthophagus subaenescens Goidanich, 1926
  Onthophagus pacificus Lansberge, 1885 China (Yunnan), N.Vietnam  
  Onthophagus panfilovi Kabakov China (Yunnan)  
  Onthophagus penicillatus Harold, 1879 China (Guizhou, Sichuan, Yunnan), Burma, Laos, Thailand, Malaysia  
  Onthophagus pentaceros Olsouffiev ,1900 S.Turkmenistan, Iran, Afganistan  
  Onthophagus poeophagus Kabakov,1979 N.E. China  
  Onthophagus ponticus Harold, 1883 S. Russia, Georgia, Azerbaidzhan, Armenia,
S.E. Europe, Turkey, N.Iran, Syria		  
  Onthophagus popovi Kabakov,1994 China (Yunnan), Vietnam  
  Onthophagus potanini Kabakov, 1979 China (Sichuan, Gansu)  
  Onthophagus procurvus Balthasar, 1935 China (Shanxi, Sichuan, Yunnan), India  
  Onthophagus productus Arrow, 1907 China (Sichuan, Yunnan), India, Bangladesh  
  Onthophagus proletarius Harold, 1875 China (Fujian, Hongkong, Hainan), Vietnam, Laos,
Taiwan, Burma, Malaysia		  
  Onthophagus proximus Kirschenblatt, China  
  Onthophagus pseudobrutus Kabakov,1994 China (Yunnan), N.Vietnam, Thailand  
  Onthophagus pseudocaccobius Reitter , 1888 Turkmenistan, Uzbekistan, Tadzhikistan  
  Onthophagus pseudojaponicus Balthasar, 1941 China Onthophagus pseudojaponicus ab. ylingensis
Balthasar, 1941
  Onthophagus pseudojaponicus ssp. fukiensis
Balthasar, 1942		 China (Fujian) Onthophagus pseudojaponicus fukiensis ab.
tschungsengi Balthasar
  Onthophagus punctator Reitter, 1893 Russia(Siberia), China (Jilin, Beijing,Shanxi), Korea,
Mongolia		 Onthophagus miyabei Matsumura, 1937;
Onthophagus sunanicus Stebnicka, 1973
  Onthophagus purpurascens Boucomont, 1914 China (Yunnan), N.Vietnam  
  Onthophagus putealis Matsumura, 1938 Taiwan  
  Onthophagus pygargus Motschulsky, 1845 Armenia, Middle Asia, Iran, Afganistan, China
(Xinjiang)		 Onthophagus saiga Ballion, 1870
  Onthophagus quadridentatus Fabriciiis, 1798 Taiwan, India, Sri-Lanka Onthophagus quadricornis Fabricius, 1801
  Onthophagus quadrinodus Reitter, 1896 Uzbekistan, Tadzhikistan  
  Onthophagus rachelis Martin, China (Yunnan, Xizang), India , Sikkim, Kashmir  
  Onthophagus rechingeri Petrovitz, 1980 S.Turkmenistan, Iran, Afganistan Onthophagus kryzhanovskii Kabakov , 1982
  Onthophagus rectecornutus Lansberge, 1883 China (Fujian, Guangdong, Guangxi, Yunnan), India,
Burma, Laos, Vietnam, Cambodia, Sri-Lanka, Bhutan,
S. Asia, Malaysia, , Indonesia		  
  Onthophagus roubali Balthasar, 1935 NE. China, Korea  
  Onthophagus rubricollis Hope, 1831 China (Yunnan,Xizang), India, Sikkim, Kashmir,
Taiwan, Burma		  
  Onthophagus rudis Sharp, 1875 China, Vietnam, Burma, India, Indonesia, Malaysia,
Thailand, Calemantan		  
  Onthophagus ruficapilllus Brulle , 1832 Russia , Ukraina, Europe, Azerbaidzhan, Armenia,
Georgia, Turkey, Turkmenistan, Irak, Syria, Iran		  
  Onthophagus rufimanus Kabakov , 1982 Turkmenistan  
  Onthophagus rugulosus Harold, 1886 Russia(Far East), China (Fujian, Sichuan, Yunnan),
Vietnam, India, Taiwan, Korea		 Onthophagus expansicollis Fairmaire, 1819
  Onthophagus rutilans Sharp, 1875 China (Fujian, Sichuan, Yunnan), Malaysia, Laos,
Korea 		 
  Onthophagus sacharovskii Olsoufflev, 1918 Georgia, Azerbaidzhan, Armenia,Turkey  
  Onthophagus sagittarius Fabricius, 1775 China (Fujian), S. China; India, Cambodia, Vietnam,
Laos, Burma, Taiwan, Indonesia		 Onthophagus oryx Fabricius; Onthophagus
japanus Fabricius, Onthophagus obtusus
Wiedeman, Onthophagus erectus Wiedemann
  Onthophagus sauteri Gillet,1924 Taiwan  
  Onthophagus scabriusculus Harold, 1873 Russia (Siberia), China (Inner Mongolia), Mongolia  
  Onthophagus schafernai Baltliasar, 1935 China  
  Onthophagus schanabeli Splichal, 1910 China (Xizang), Kashmir , Afganistan  
  Onthophagus semicornis Panzer , 1798 Russia, Belorussia, Turkmenistan, Kazahstan,
Georgia, Azerbaidzhan, Armenia, Europe, Turkey,
N. Iran		 angulicornis ,- decipiens Mulsant, 1842
  Onthophagus senex Boucomont, 1914 China (Guangzhou, Hongkong, Sichuan), India,
Burma, Laos		  
  Onthophagus seniculus Fabricius, 1781 China (Guangzhou, Hongkong, Guangxi, Yunnan),
India, Burma, Cambodia, Taiwan, Thailand, Malaysia		  
  Onthophagus sericatus Reitter , 1893 S. Russia (Caucasus), S.E. Europe, Georgia,
Azerbaidzhan, Armenia, Lebanon , Iran		  
  Onthophagus setchan Masumoto, 1984 China  
  Onthophagus sibiricus Harold, 1877 Turkmenistan, Kazahstan, Uzbekistan, Kirgizstan,
Tadzhikistan, Mongolia, N. China		  
  Onthophagus silus Balthasar , 1960 Turkmenistan, Kazahstan,Uzbekistan, Tadzhikistan Onthophagus kirgisicus Protzenko , 1963
  Onthophagus similis Scriba , 1790 Europe, W.Belorussia, W. Ukraina  
  Onthophagus smetannai Balthasar, 1952 China (Fujian)  
  Onthophagus sobrius Balthasar, 1960 China  
  Onthophagus solivagus Harold, 1886 Russia (Far East), China (Beijing, Shanxi, Hubei,
Sichuan, Gansu), Japan, Korea		  
  Onthophagus sonai Miwa, 1930 Taiwan  
  Onthophagus speculifer Solsky, 1876 Kazahstan, Kirgizstan, Turkmenistan, Uzbekistan,
E. Iran, Afganistan		  
  Onthophagus sternax Balthasar, 1960 S. China  
  Onthophagus strandi Balthasar, 1942 China (Fujian)  
  Onthophagus suermelii Petrovitz , 1963 Armenia, Turkey  
  Onthophagus sulci Balthasar, 1935 China (Fujian, Sichuan)  
  Onthophagus sulcicollis Reitter ,1892 Kazahstan, Turkmenistan, Uzbekistan, Kirgizstan,
Iran, Afganistan 		 
  Onthophagus susterai Balthasar, 1952 China  
  Onthophagus sutleinensis Splichal, 1910 China (Xizang), N.India Onthophagus concursator Balthasar, 1960
  Onthophagus suturellus Brulle , 1832 Russia , Ukraina, Georgia, Azerbaidzhan, Armenia,
Turkey, S.E. Europe, S.Turkmenistan, N. Iran		 Onthophagus trochiscobius Kolenati, 1846
  Onthophagus sycophanta Fairmaire, 1887 China (Fujian, Hunan,Guizhou, Sichuan , Yunnan),
Vietnam		  
  Onthophagus tabidus Balthasar, 1935 China  
  Onthophagus taiwanus Nomura, Taiwan  
  Onthophagus taiyaruensis Masumoto, 1977 Taiwan  
  Onthophagus tatsienluensis Balthasar, 1942 China (Fujian, Sichuan)  
  Onthophagus taurinus White, 1844 China(Hubei, Shanghai, Fujian), S. China (Hongkong),
India, Burma, Taiwan, Laos, Vietnam, Cambodia,
Thailand, Bangladesh 		 
  Onthophagus taurus Schreber , 1759 Russia , Ukraina, Europe, N. Afrika, Turkey,
Azerbaidzhan, Armenia, Georgia, Kazahstan,
Turkmenistan, Kazahstan, Uzbekistan, Kirgizstan,
Tadzhikistan, Afganistan, Iran ,N. China		 Onthophagus rugosus Poda, 1761;corniger
Geoffroy 1785; recticornis Leske,1785; capra
Fabricius, 1787
  Onthophagus tibetanus Arrow, 1907 China (Xizang)  
  Onthophagus tragus Fabricius, 1792 China (Beijing, Shanxi, Fujiari, Guangdong,
Hongkong, Yunnan), India, Taiwan, Burma, Vietnam,
Bangladesh, Malaysia, Indonesia, Celebes		 Onthophagus fulvicornis Motschulsky, 1853;
Onthophagus mandarinus Harold, 1859;
Onthophagus ibex Motschulsky, 1854
  Onthophagus tragoides Boucomont, 1914 China (Yunnan), Vietnam  
  Onthophagus transcaspicus Koenig , 1888 S. Turkmenistan, S. Uzbekistan, Iran, Afganistan,
Pakistan, N. Afrika		 Onthophagus lineatus Reitter, 1893;
Onthophagus gasella Heyden, 1913
  Onthophagus tricolor Boucomont, 1914 China (Fujian, Guangdong), Vietnam, Indonesia  
  Onthophagus tricornis Wiedeman, 1823 China (Henan, Hubei, Shanghai, Guarigxi), India,
Burma, Taiwan, Cambodia, Vietnam , Laos,
Indonesia, Malaysia, Japan		 Onthophagus rectinicornis Fairmaire, 1878
  Onthophagus trispinus Reitter, 1892 Tadzhikistan, S.Turkmenistan, Uzbekistan,
N. Afganistan 		Onthophagus umbilicopunctatus Medvedev,
1959
  Onthophagus tritinctus Boucomont, 1914 China (Hongkong), India , Thailand, Sri-Lanka  
  Onthophagus trituber trituber Wiedeman, 1823 China (Shanghai, Guangdong, Hongkong), Vietnam,
Laos, Taiwan, Malaysia, Indonesia		 Onthophagus suturalis White, 1844
  Onthophagus trituber jakobsoni Kabakov, 2006 N.E. China  
  Onthophagus truchmenus truchmenus Kolenati
, 1846		 Georgia, Azerbaidzhan, Armenia, Turkmenistan, Iran,
Syria		  
  Onthophagus truchmenus iranicus Kabakov,
2006 		S.Iran  
  Onthophagus truchmenus turkmenicus Kabakov,2006 Turkmenistan  
  Onthophagus turfanicus Balthasar, China(Fujian)  
  Onthophagus turmalis Gillet,1924 Taiwan  
  Onthophagus turpidus Reitter, 1887 China(Xinjiang, Sichuan, Xizang)  
  Onthophagus turpidoides Kabakov China (Gansu, Sichuan)  
  Onthophagus umenoi Matsumura, 1938 Taiwan  
  Onthophagus unguiculatus Kabakov,1994 China (Yunnan)  
  Onthophagus uniformis Heyden, 1886 Russia(Far East); China (Gansu, Beijing), Korea  
  Onthophagus vacca Linnaeus ,1767 Russia ,Ukraina, Kazahstan, Turkmenistan, Europe,
N. Afrika, Iran, Syria		 Onthophagus conspurcatus Geoffroy, 1785 ;
Onthophagus medius Kugelan, 1792; Onthophagus
aeruginosus Schrank,1798 ; Onthophagus affinis
Sturm , 1800; Onthophagus tricornis Fischer -
Waldh, 1844; Onthophagus antilope
Motschulsky, 1845; Onthophagus vicinus, -
difficilis, - intermedius , - propinquus, - similes,
- sublineolatus,- basalis Mulsant, 1842
  Onthophagus vaulogeri Boucomont, 1923 China (Yunnan), N.Vietnam  
  Onthophagus verticicornis Laicharting , 1781 S. Russia, Ukraina, Europe Onthophagus nutans Fabricius , 1787 ;
Onthophagus distinguendus,- infuscatus
Mulsant, 1842
  Onthophagus viduus Harold, 1874 China (Sichuan, Beijing, Hubei, Jiangxi, Shanghai,
Guangxi) Japan, Korea		 Onthophagus flavonotatus d‘Orbigny, 1898
  Onthophagus vigilans Boucomont, 1921 China (Yunnan), Vietnam  
  Onthophagus viridis Ménétriès, 1832 Azerbaidzhan, N. Iran, S. Turkmenistan  
  Onthophagus vitulus Fabricius , 1776 Russia, Moldavia, Ukraina, Europe, Caucasus , Iran,
Kazahstan, Turkmenistan, Uzbekistan,Kirgizstan,
N. China		 Onthophagus camelus Fabricius , 1777;
Onthophagus kaszabi Balthasar, 1963
  Onthophagus vlasovi Medvedev , 1958 Turkmenistan, Kazahstan, Uzbekistan, Kirgizstan,
Tadzhikistan, N.E. Iran , Afganistan 		 
  Onthophagus (Micronthpagus) wangi
Masumoto et Chen et Ochi, 2004		    
  Onthophagus yanoi Matsumura, 1938 Taiwan  
  Onthophagus yubarinus Matsumura, 1937 Taiwan  
  Onthophagus yunnanus Boucomont, 1912 China ( Hubei, Fujian,Guizhou, Sichuan, Yunnan)  
  Onthophagus zavreli Balthasar, 1935 China( Hubei, Sichuan,Yunnan)  
  Onthophagus zimmermanni Balthasar, 1960 China( Sichuan)  
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